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THE LOST TYPES OF ROTALIA BECCARII VAR. TEPIDA CUSHMAN 1926

¹ Department of Geology, University of Auckland, Private Bag 92019, Auckland, New Zealand. Email: b.hayward{at}geomarine.org.nz
² Department of Paleobiology, NMNH, Smithsonian Institution, Washington D.C., 20560, USA.
³ Department of Biology, Midwestern State University, 3410 Taft Blvd., Wichita Falls, Texas, 76308, USA.
⁴ Station de Zoologie, 154, Université de Genève, Route de Malagnou 154, 1224 Chêne-Bougeries, Switzerland.

In 1926, Cushman described Rotalia beccarii var. tepida from San Juan Harbor, Puerto Rico. His publication illustrates a strew slide with many individuals, and no holotype was designated. Consequently, over the years, researchers have assumed that no holotype exists. A search of the Cushman Collection, however, discovered a slide labeled as holotype and it is so recorded in the Cushman Catalog of 1929. Here, this specimen is re-described and designated as a lectotype. All nine of the other surviving syntypes from the original collection in San Juan Harbor, Puerto Rico, now become paralectotypes. Hopefully, the designation of formal types for Ammonia tepida (Cushman) will help stabilize the taxonomy of this important species. Our study shows that the lectotype is morphologically distinguishable from other Ammonia types and that the distribution of Ammonia tepida is restricted to tropical shallow-water environments.

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